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What is Forretress based on?

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Its evolution, Pineco, is clearly based off a pinecone, but Forretress shares no resemblance with anything I can think of.

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Forretress is based off of a bagworm, that is possibly inhabiting a discarded shell of some kind, and with she spikes on it's shell, it could also be based on mussels, shellfish, or a giant clam. Hope this helped!

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Check at species. It says on it's page that is a bagworm: http://pokemondb.net/pokedex/forretress.