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I know it removes the flying type, but certain non-flying but levitating pokemon can learn it. (such as Flygon)

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Umm, Swaggron, I'm talking about Levitating pokemon, not Flying types.
Well its kind of the same

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No, they will not be vulnerable to Ground Type Moves.

If a non-Flying-type uses Roost, then there are no side-effects.

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Thank you.