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When you're trying to catch the lake trio in Nameless Cavern, (during the day) do Dusk Balls still get their bonus for being in a cave? Or since you battle in the portal-area, does that make it not in the cave anymore?

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Yes, they do. The game acts as if the battle is in the area, despite it in the ring.(source:expirence)

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Ok, ima go catch the lake trio now. thanks!