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On this site it says only in gen4, but other sites state that 25% of the time in rain, Thunder can break through Protect/Detect. Is this true?

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No
Thunder had a 30% chance to hit through Protect in Diamond, Pearl, Heartgold, and Soul Silver. From Platinum on Thunder will not Hit through protect.

Source: http://bulbapedia.bulbagarden.net/wiki/Thunder_(move)

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